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Probatova. 2005. A progress in the study of the chromosome numbers of rare and endemic species of the flora of the Russian Far East. Pages 85--86 in Karyology, Karyosystematics and Molecular Phylogeny. St. Petersburg, Russia.
Probatova. 2004. Chromosome numbers of selected vascular plant species from Sakhalin, Moneron and the Kurile Islands. Biodivers. Biogeogr. Kuril Islands Sakhalin 1: 15–23.